"Some men probably did inherit it, but then consequently tended to reproduce less than the other men who did not inherit it. In order for a trait to become established, not only does it have to be inherited, but it has to be inherited by those who then reproduced more than (or at least not less than) average."
But by the same token, how did gracility for women become established? Since the female offspring also have the potential of patrilineally inheriting their skeletal type, wouldn't there have been a sizeable proportion of robust women as well? And wouldn't these women have reproduced as much as the gracile women, since gracility does not afford any ergonomic advantages after the women have been transported back to the steppe?